On the Origin of Species by Charles Darwin (grave mercy .txt) 📕

>Hence it has come, that when we compare the now living productions of the temperate regions of the New and Old Worlds, we find very few identical species (though Asa Gray has lately shown that more plants are identical than was formerly supposed), but we find in every great class many forms, which some naturalists rank as geographical races, and others as distinct species; and a host of closely allied or representative forms which are ranked by all naturalists as specifically distinct.

 

As on the land, so in the waters of the sea, a slow southern migration of a marine fauna, which during the Pliocene or even a somewhat earlier period, was nearly uniform along the continuous shores of the Polar Circle, will account, on the theory of modification, for many closely allied forms now living in areas completely sundered. Thus, I think, we can understand the presence of many existing and tertiary representative forms on the eastern and western shores of temperate North America; and the still more striking case of many closely allied crustaceans (as described in Dana’s admirable work), of some fish and other marine animals, in the Mediterranean and in the seas of Japan,—areas now separated by a continent and by nearly a hemisphere of equatorial ocean.

 

These cases of relationship, without identity, of the inhabitants of seas now disjoined, and likewise of the past and present inhabitants of the temperate lands of North America and Europe, are inexplicable on the theory of creation. We cannot say that they have been created alike, in correspondence with the nearly similar physical conditions of the areas; for if we compare, for instance, certain parts of South America with the southern continents of the Old World, we see countries closely corresponding in all their physical conditions, but with their inhabitants utterly dissimilar.

 

But we must return to our more immediate subject, the Glacial period.

I am convinced that Forbes’s view may be largely extended. In Europe we have the plainest evidence of the cold period, from the western shores of Britain to the Oural range, and southward to the Pyrenees.

We may infer, from the frozen mammals and nature of the mountain vegetation, that Siberia was similarly affected. Along the Himalaya, at points 900 miles apart, glaciers have left the marks of their former low descent; and in Sikkim, Dr. Hooker saw maize growing on gigantic ancient moraines. South of the equator, we have some direct evidence of former glacial action in New Zealand; and the same plants, found on widely separated mountains in this island, tell the same story. If one account which has been published can be trusted, we have direct evidence of glacial action in the south-eastern corner of Australia.

 

Looking to America; in the northern half, ice-borne fragments of rock have been observed on the eastern side as far south as lat. 36 deg-37

deg, and on the shores of the Pacific, where the climate is now so different, as far south as lat. 46 deg; erratic boulders have, also, been noticed on the Rocky Mountains. In the Cordillera of Equatorial South America, glaciers once extended far below their present level.

In central Chile I was astonished at the structure of a vast mound of detritus, about 800 feet in height, crossing a valley of the Andes; and this I now feel convinced was a gigantic moraine, left far below any existing glacier. Further south on both sides of the continent, from lat. 41 deg to the southernmost extremity, we have the clearest evidence of former glacial action, in huge boulders transported far from their parent source.

 

We do not know that the Glacial epoch was strictly simultaneous at these several far distant points on opposite sides of the world. But we have good evidence in almost every case, that the epoch was included within the latest geological period. We have, also, excellent evidence, that it endured for an enormous time, as measured by years, at each point. The cold may have come on, or have ceased, earlier at one point of the globe than at another, but seeing that it endured for long at each, and that it was contemporaneous in a geological sense, it seems to me probable that it was, during a part at least of the period, actually simultaneous throughout the world. Without some distinct evidence to the contrary, we may at least admit as probable that the glacial action was simultaneous on the eastern and western sides of North America, in the Cordillera under the equator and under the warmer temperate zones, and on both sides of the southern extremity of the continent. If this be admitted, it is difficult to avoid believing that the temperature of the whole world was at this period simultaneously cooler. But it would suffice for my purpose, if the temperature was at the same time lower along certain broad belts of longitude.

 

On this view of the whole world, or at least of broad longitudinal belts, having been simultaneously colder from pole to pole, much light can be thrown on the present distribution of identical and allied species. In America, Dr. Hooker has shown that between forty and fifty of the flowering plants of Tierra del Fuego, forming no inconsiderable part of its scanty flora, are common to Europe, enormously remote as these two points are; and there are many closely allied species. On the lofty mountains of equatorial America a host of peculiar species belonging to European genera occur. On the highest mountains of Brazil, some few European genera were found by Gardner, which do not exist in the wide intervening hot countries. So on the Silla of Caraccas the illustrious Humboldt long ago found species belonging to genera characteristic of the Cordillera. On the mountains of Abyssinia, several European forms and some few representatives of the peculiar flora of the Cape of Good Hope occur. At the Cape of Good Hope a very few European species, believed not to have been introduced by man, and on the mountains, some few representative European forms are found, which have not been discovered in the intertropical parts of Africa. On the Himalaya, and on the isolated mountain-ranges of the peninsula of India, on the heights of Ceylon, and on the volcanic cones of Java, many plants occur, either identically the same or representing each other, and at the same time representing plants of Europe, not found in the intervening hot lowlands. A list of the genera collected on the loftier peaks of Java raises a picture of a collection made on a hill in Europe! Still more striking is the fact that southern Australian forms are clearly represented by plants growing on the summits of the mountains of Borneo. Some of these Australian forms, as I hear from Dr. Hooker, extend along the heights of the peninsula of Malacca, and are thinly scattered, on the one hand over India and on the other as far north as Japan.

 

On the southern mountains of Australia, Dr. F. Muller has discovered several European species; other species, not introduced by man, occur on the lowlands; and a long list can be given, as I am informed by Dr.

Hooker, of European genera, found in Australia, but not in the intermediate torrid regions. In the admirable ‘Introduction to the Flora of New Zealand,’ by Dr. Hooker, analogous and striking facts are given in regard to the plants of that large island. Hence we see that throughout the world, the plants growing on the more lofty mountains, and on the temperate lowlands of the northern and southern hemispheres, are sometimes identically the same; but they are much oftener specifically distinct, though related to each other in a most remarkable manner.

 

This brief abstract applies to plants alone: some strictly analogous facts could be given on the distribution of terrestrial animals. In marine productions, similar cases occur; as an example, I may quote a remark by the highest authority, Professor Dana, that “it is certainly a wonderful fact that New Zealand should have a closer resemblance in its crustacea to Great Britain, its antipode, than to any other part of the world.” Sir J. Richardson, also, speaks of the reappearance on the shores of New Zealand, Tasmania, etc., of northern forms of fish.

Dr. Hooker informs me that twenty-five species of Algae are common to New Zealand and to Europe, but have not been found in the intermediate tropical seas.

 

It should be observed that the northern species and forms found in the southern parts of the southern hemisphere, and on the mountain-ranges of the intertropical regions, are not arctic, but belong to the northern temperate zones. As Mr. H. C. Watson has recently remarked, “In receding from polar towards equatorial latitudes, the Alpine or mountain floras really become less and less arctic.” Many of the forms living on the mountains of the warmer regions of the earth and in the southern hemisphere are of doubtful value, being ranked by some naturalists as specifically distinct, by others as varieties; but some are certainly identical, and many, though closely related to northern forms, must be ranked as distinct species.

 

Now let us see what light can be thrown on the foregoing facts, on the belief, supported as it is by a large body of geological evidence, that the whole world, or a large part of it, was during the Glacial period simultaneously much colder than at present. The Glacial period, as measured by years, must have been very long; and when we remember over what vast spaces some naturalised plants and animals have spread within a few centuries, this period will have been ample for any amount of migration. As the cold came slowly on, all the tropical plants and other productions will have retreated from both sides towards the equator, followed in the rear by the temperate productions, and these by the arctic; but with the latter we are not now concerned. The tropical plants probably suffered much extinction; how much no one can say; perhaps formerly the tropics supported as many species as we see at the present day crowded together at the Cape of Good Hope, and in parts of temperate Australia. As we know that many tropical plants and animals can withstand a considerable amount of cold, many might have escaped extermination during a moderate fall of temperature, more especially by escaping into the warmest spots.

But the great fact to bear in mind is, that all tropical productions will have suffered to a certain extent. On the other hand, the temperate productions, after migrating nearer to the equator, though they will have been placed under somewhat new conditions, will have suffered less. And it is certain that many temperate plants, if protected from the inroads of competitors, can withstand a much warmer climate than their own. Hence, it seems to me possible, bearing in mind that the tropical productions were in a suffering state and could not have presented a firm front against intruders, that a certain number of the more vigorous and dominant temperate forms might have penetrated the native ranks and have reached or even crossed the equator. The invasion would, of course, have been greatly favoured by high land, and perhaps by a dry climate; for Dr. Falconer informs me that it is the damp with the heat of the tropics which is so destructive to perennial plants from a temperate climate. On the other hand, the most humid and hottest districts will have afforded an asylum to the tropical natives. The mountain-ranges north-west of the Himalaya, and the long line of the Cordillera, seem to have afforded two great lines of invasion: and it is a striking fact, lately communicated to me by Dr. Hooker, that all the flowering plants, about forty-six in number, common to Tierra del Fuego and to Europe still exist in North America, which must have lain on the