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by the branching of a sensory axon. (Figure text: cord, sensory neurone, motor neurone)

Thus the nerve current from a single sensory neurone is distributed to quite a number of motor neurones. Where there are central neurones in the arc, their branching axons aid in distributing the excitation; and so we get a big movement in response to a minute, though intense stimulus.

But the response is not simply big; it is definite, coordinated, representing team work on the part of the muscles as distinguished from indiscriminate mass action. That means selective distribution of the nerve current. The axons of the sensory and central neurones do not connect with any and every motor neurone indiscriminately, but link up with selected groups of motor neurones, and thus harness together teams that will work in definite ways, producing {39} flexion of a limb in the case of one such team, and extension in the case of another. Every reflex has its own team of motor neurones, harnessed together by its outfit of sensory and central neurones. The same motor neurone may however be harnessed into two or more such teams, as is seen from the fact that the same muscle may participate in different reflex movements; and for a similar reason we believe that the same sensory neurone may be utilized in more than one reflex arc.



Fig. 11.--Coรถrdination brought about by the branching of the axon of a central neurone. (Figure text: sensory, central, motor)

The most distinctive part of any reflex arc is likely to be its central neurones, which are believed to play the chief part in coรถrdination, and in determining the peculiarities of any given reflex, such as its speed and rhythm of action.

Reactions in General

Though the reflex is simple by comparison with voluntary movements, it is not the simplest animal reaction, for it is coรถrdinated and depends on the nervous system, while the simplest animals, one-celled animals, have no nervous system, any more than they have muscles or organs of any {40} kind. Without possessing separate organs for the different vital functions, these little creatures do nevertheless take in and digest food, reproduce their kind, and move. Every animal shows at least two different motor reactions, a positive or approaching reaction, and a negative or avoiding reaction.

The general notion of a reaction is that of a response to a stimulus. The stimulus acts on the organism and the organism acts back. If I am struck by a wave and rolled over on the beach, that is passive motion and not my reaction; but if the wave stimulates me to maintain my footing, then I am active, I respond or react.

Now there is no such thing as wholly passive motion. Did not Newton teach that "action and reaction are equal"?--and he was thinking of stones and other inanimate objects. The motion of a stone or ball depends on its own weight and shape and elasticity as much as on the blow it receives. Even the stone counts for something in determining its own behavior.

A loaded gun counts for more than a stone, because of the stored energy of the powder that is set free by the blow of the hammer. The "reaction" of the gun is greater than the force acting on it, because of this stored energy that is discharged.

An animal reaction resembles the discharge of the gun, since there is stored energy in the animal, consisting in the chemical attraction between food absorbed and oxygen inspired, and some of this energy is utilized and converted into motion when the animal reacts. The stimulus, like the trigger of the gun, simply releases this stored energy.

The organism, animal or human, fully obeys the law of conservation of energy, all the energy it puts out being accounted for by stored energy it has taken in in food and oxygen. But at any one time, when the organism receives {41} a stimulus, the energy that it puts forth in reaction comes from inside itself.

There is another way in which the organism counts in determining its reaction. Not only does it supply the energy of the response, but its own internal arrangements determine how that energy shall be directed. That is to say, the organism does not blow up indiscriminately, like a charge of dynamite, but makes some definite movement. This is true even of the simplest animals, and the more elaborate the internal mechanism of the animal, the more the animal itself has to do with the kind of response it shall make to a stimulus. The nervous system of the higher animals, by the connections it provides between the stimulus and the stores of energy in the muscles, is of especial importance in determining the nature of the response.

Stimuli are necessary to arouse the activity of the organism. Without any stimulus whatever, it seems likely that the animal would relapse into total inactivity. It should be said, however, that stimuli, such as that of hunger, may arise within the organism itself. The stimulus may be external or internal, but some stimulus is necessary in order to release the stored energy.

In general, then, a reaction consists in the release by a stimulus of some of the stored energy of an animal, and the direction of that energy by the animal's own internal mechanism of nerves and muscles (and, we may add, bones and sinews) into the form of some definite response.

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EXERCISES

1. Outline of the chapter, being at the same time a "completion test". Complete the following outline by filling in the blank spaces (usually a single word will fill the blank, but sometimes two words will be better):

A. Definition: A reaction is a response to a ___________. The stimulus energy stored in the organism, and the __________ has a definite form determined by the organism's own machinery of ________ and ______. 

B. Among very prompt reactions are the reflex and the "simple reaction". The reflex differs from the "simple reaction" in that:

(1) It usually takes less________.

(2) It requires no___________,

(3) The machinery for it is ________in the organism.

C. The machinery for a reflex consists of:

(1) a________ organ.

(2) a ________nerve.

(3) a nerve ________,

(4) a _________nerve.

(5) a muscle or _________.

D. The sensory and motor nerves consist of ________ which are branches of ______. The cells for the motor nerves lie in the ________, and those for the sensory nerves lie in two cases in the _________, and in all other cases in bunches located close beside the _________or ________,

E. The neurone is the _______ of which the nervous ______ is composed. It consists of a ________ and of two sorts of branches, the ________ and the ________. Internally, the neurone shows a peculiar structure of ________ and ________.

F. Communication from one neurone to another occurs across a _____ called the synapse. The _________of an axon here comes into close contact with the ______or with the _________of another neurone. The communication takes place from the ________of the first neurone to the ___________ of the second.

G. The "nerve current" in a reflex therefore runs the following course: from the sense organ into a ________ axon, along this to its _________ in a nerve, and across a _________ there into the _________ of a neurone, and thence {43} out along the _______of this neurone to the ________or _________ that executes the reflex. This is a two-neurone _________, but often there is a third, ________neurone between the _________ and the _____________.

H. Coรถrdination is effected by the ________ of the axons of the sensory and ________ neurones, by which means the nerve current is ______ to a team of ________ and so to a team of _________.

2. Is the reaction time experiment, as described in the text, an introspective or an objective experiment?

3. Mention two cases from common life that belong under the "simple reaction", two that belong under "choice reaction", and two that belong under the "associative reaction".

4. Arrange the reflexes mentioned in the text under the two heads of "protective" and "regulative".

5. Draw diagrams of (a) the neurone, (b) a synapse, (c) a reflex arc, and (d) a coรถrdinated movement. Reduce each drawing to the simplest possible form, and still retain everything that is essential.

6. What part of the nervous system lies (a) in the forehead and top of the head, (b) in the very back of the head, (c) along the base of the skull, (d) within the backbone, (e) in the arm?

7. Using a watch to take the time, see how long it takes you to name the letters in a line of print, reading them in reverse order from the end of the line to the beginning. Compare with this time the time required to respond to each letter by the letter following it in the alphabet (saying "n" when you see m, and "t" when you see s, etc.). Which of these two "stunts" is more like reflex action, and how, nevertheless, does it differ from true reflex action?

8. The pupillary reflex. Describe the reaction of the pupil of the eye to light suddenly shining into the eye. This response can best be observed in another person, but you can observe it in yourself by aid of a hand mirror. On another person you can also observe the "crossed" pupillary reflex, by throwing the light into one eye only while you watch the other eye. What sort of connection do you suppose to exist between the two eyes, making this crossed reflex possible?

9. The lid reflex, or wink reflex, (a) Bring your hand suddenly close to another person's eye, and notice the response of the eyelid, (b) See whether you can get a crossed reflex here, (c) See whether your subject can voluntarily prevent (inhibit) the lid reflex, (d) See whether the reflex occurs when he gives the stimulus himself, by moving his own hand suddenly up to his eye. (e) What other stimulus, besides the visual one that you have been using, will arouse the same response?

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REFERENCES

C. Judson Herrick, in his Introduction to Neurology, 2nd edition, 1918, gives a fuller and yet not too detailed account of the neurone in Chapter III, and of reflex action in Chapter IV.

Percy G. Stiles, in his Nervous System and Its Conservation, 1915, discusses these matters in Chapters II, III and IV.

Ladd and Woodworth's Elements of Physiological Psychology, 1911, has chapters on these topics.

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CHAPTER III
REACTIONS OF DIFFERENT LEVELS


HOW SENSATIONS, PERCEPTIONS AND THOUGHTS MAY BE CONSIDERED AS FORMS OF INNER RESPONSE, AND HOW THESE HIGHER REACTIONS ARE RELATED IN THE NERVOUS SYSTEM TO THE SIMPLER RESPONSES OF THE REFLEX LEVEL.

Having defined a reaction as an act of the individual aroused by a stimulus, there is no reason why we should not include a great variety of mental processes under the general head of reactions. Any mental process is an activity of the organism, and it is aroused by some stimulus, external or internal; therefore, it is a reaction.

I hear a noise--now, while the noise, as a physical stimulus, comes to me, my hearing it is my own act, my sensory reaction to the stimulus. I recognize the noise as the whistle of a steamboat--this recognition is clearly my own doing, dependent on my own past experience, and may be called a perception or perceptive response. The boat's whistle reminds me of a vacation spent on an island--clearly a memory response. The memory arouses an agreeable feeling--an affective response, this may be called. In its turn, this may lead me to imagine how pleasant it would be to spend another vacation on that island, and to cast about for ways and means to accomplish this result--here we have imagination and reasoning, aroused by what preceded just as the sensation was aroused by the physical stimulus.

In speaking of any mental process as an act of the individual, we do not mean to imply that he is always conscious {46} of his activity. Sometimes he feels active, sometimes passive. He feels

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